The llama (Lama glama) is a South American camelid, widely used as a pack animal by the Incas and other natives of the Andes mountains. In South America and some parts of Saudi Arabia llamas are still used for beasts of burden, fiber production and meat.
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The height of a full-grown, full-size llama is between 5.5
feet (1.6 meters) to 6 feet (1.8 meters) tall at the top of the
head. They can weigh approximately between 280 pounds (127
kilograms) and 450 pounds (204 kilograms). At birth, a baby
llama (called a cria) can weigh between 20 pounds (9 kilograms)
to 30 pounds (14 kilograms). Llamas are very social animals and
like to live with other llamas as a herd. Overall, the fiber
produced by a llama is very soft and is naturally lanolin free.
Very intelligent, llamas learn simple tasks after a few
repetitions. When using a pack, llamas can carry about 25% - 30%
of their body weight for several miles.
Llamas originated from the central plains of North America about
40 million years ago. They migrated to South America and Asia
about 3 million years ago. By the end of the last ice-age
(10,000 - 12,000 years ago) camelids were extinct in North
America. As of 2007, there were over 7 million llamas and
alpacas in South America and due to importation from South
America in the late 20th Century there are now over 100,000
llamas and 6,500 - 7,000 alpacas in the US and Canada.
Although they were often compared by early writers to sheep and
spoken of as such, their similarity to the camel was very soon
perceived. They were included in the genus Camelus in the
Systema Naturae of Linnaeus. They were, however, separated by
Cuvier in 1800 under the name of llama along with the alpaca and
the guanaco. Vicuñas are in genus Vicugna. The animals of the
genus Lama are, with the two species of true camels, the sole
existing representatives of a very distinct section of the "Artiodactyla"
or even-toed ungulates, called Tylopoda, or "bump-footed," from
the peculiar bumps on the soles of their feet, on which they
tread. This section thus consists of a single family, the
Camelidae, the other sections of the same great division being
the Suina or pigs, the Tragulina or chevrotains, and the Pecora
or true ruminants, to each of which the Tylopoda have more or
less affinity, standing in some respects in a central position
between them, borrowing as it were some characters from each,
but in others showing great special modifications not found in
any of the other sections.
The discoveries of a vast and previously unsuspected extinct
fauna of the American continent of the Tertiary period, as
interpreted by the palaeontologists Leidy, Cope, and Marsh, has
thrown a flood of light upon the early history of this family,
and upon its relations to other mammals. It is now known that
llamas at one time were not confined to the part of the
continent south of the Isthmus of Panama, as at the present day,
for their remains have been abundantly found in the Pleistocene
deposits of the region of the Rocky Mountains, and in Central
America, some attaining a much larger size than those now
existing. Some species of llamas did stay in North America
during the last ice ages. 25,000 years ago, llamas would have
been a common sight in modern-day California, Texas, New Mexico,
Utah, Missouri, and Florida. These North American llamas belong
to a single genus, Hemiauchenia, which is extinct.
Many camel-like animals exhibiting different genetic
modifications and a gradual series of changes, coinciding with
the antiquity of the deposits in which they are found, have been
traced from the thoroughly differentiated species of the modern
epoch down through the Pliocene to the early Miocene beds. Their
characters having become more generalized, they have lost all
that especially distinguishes them as Camelidae: they are merged
into forms common to the ancestral type of all the other
sections of the Artiodactyles.
So far none of these annectant forms have been found in any of
the fossiliferous strata of the Old World; it may therefore be
fairly surmised (according to the evidence at present before us)
that the Americas were the original home of the Tylopoda, and
that the true camels have passed over into the Old World,
probably by way of north Asia. Gradually driven southward,
perhaps by changes of climate, and having become isolated, they
have undergone further special modifications. Meanwhile, those
members of the family that remained in their original birthplace
have become, through causes not clearly understood, restricted
solely to the southern or most distant part of the continent.
There are few groups of mammals for which the palaeontological
history has been so satisfactorily demonstrated as the llama.
The following characteristics apply especially to llamas.
Dentition of adults:-incisors 1/3 canines 1/1, premolars 2/2,
molars 3/2; total 32. In the upper jaw there is a compressed,
sharp, pointed laniariform incisor near the hinder edge of the
premaxilla, followed in the male at least by a moderate-sized,
pointed, curved spank canine in the anterior part of the
maxilla. The isolated canine-like premolar which follows in the
camels is not present. The teeth of the molar series which are
in contact with each other consist of two very small premolars
(the first almost rudimentary) and three broad molars,
constructed generally like those of Camelus. In the lower jaw,
the three incisors are long, spatulate, and procumbent; the
outer ones are the smallest. Next to these is a curved, suberect
canine, followed after an interval by an isolated minute and
often deciduous simple conical premolar; then a contiguous
series of one premolar and three molars, which differ from those
of Camelus in having a small accessory column at the anterior
outer edge.
The skull generally resembles that of Camelus, the relatively
larger brain-cavity and orbits and less developed cranial ridges
being due to its smaller size. The nasal bones are shorter and
broader, and are joined by the premaxilla.
Vertebrae:
The ears are rather long and slightly curved inward,
characteristically known as "banana" shaped. There is no dorsal
hump. Feet are narrow, the toes being more separated than in the
camels, each having a distinct plantar pad. The tail is short,
and fibre is long, woolly and soft.
In essential structural characteristics, as well as in general
appearance and habits, all the animals of this genus very
closely resemble each other, so that whether they should be
considered as belonging to one, two, or more species is a matter
of controversy among naturalists.
The question is complicated by the circumstance of the great
majority of individuals which have come under observation being
either in a completely or partially domesticated state. Many are
also descended from ancestors which have previously been
domesticated; a state which tends to produce a certain amount of
variation from the original type. It has, however, lost much of
its importance since the doctrine of the distinct origin of
species has been generally abandoned. The four forms commonly
distinguished by the inhabitants of South America are recognized
by some naturalists as distinct species, and have had specific
designations attached to them, though usually with expressions
of doubt, and with great difficulties in defining their
distinctive characteristics.
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Llamas have an unusual reproductive cycle for a large animal.
Female llamas are induced ovulators. Through the act of mating,
the female releases an egg and is often fertilized on the first
attempt. Female llamas do not go into "heat" or have an estrus
cycle.
Like humans, llama males and females mature sexually at
different rates. Females reach puberty at approximately 12
months. However, males do not become sexually mature until
approximately 3 years.
Differentiating characteristics between llamas and alpacas
include the llama's larger size and longer head. Alpaca fiber is
generally more expensive but not always more valuable. Alpacas
tend to have a more consistent color throughout the body. The
most apparent visual difference between llamas and camels is
that camels have a hump or humps and llamas do not.
Llamas mate with the female in a kush (lying down) position,
which is fairly unusual in a large animal. They mate for an
extended period of time (20–45 minutes), also unusual in a large
animal.
The gestation period of a llama is 11 1/2 months (350 days).
Dams (female llamas) do not lick off their babies, as they have
an attached tongue which does not reach outside of the mouth
more than half an inch. Rather, they will nuzzle and hum to
their newborns.
A female is turned out into a field with a male llama and left
there for some period of time. This is the easiest method in
terms of labor, but the least useful in terms of prediction of a
likely birth date. An ultrasound test can be performed and
together with the exposure dates a better idea when the cria is
expected can be determined.
This is the most efficient method, but requires the most work on
the part of the human involved. A male and female llama are put
into the same pen and breeding is monitored. They are then
separated and rebred every other day until one or the other
refuses the breeding. Usually one can get in two breedings using
this method, though some studs have routinely refused to breed a
female more than once. The separation presumably helps to keep
the sperm count high for each breeding and also helps to keep
the condition of the female llama's reproductive tract more
sound. If the breeding is not successful within two to three
weeks, the female is rebred once again.

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